Kois, domesticated varieties of the common carp also known as Japanese carp, can live for centuries. The most famous scarlet koi, named "Hanako" (1751 - 1977), was owned by several individuals, the last of whom was Dr. Komei Koshihara. Hanako was reportedly 226 years old upon her death; her age was determined by removing one of her scales and examining it extensively in 1966.
She is, to date, the longest-lived vertebrate ever recorded.
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